An international terrorist for hire named Castor Troy is being relentlessly pursued by Sean Archer, an FBI agent whose son was killed years earlier when Troy attempted to kill Archer. The FBI receives information that Castor's brother Pollux has chartered a plane at a Los Angeles airport, leading Archer to believe that Castor is with him. Archer leads an FBI team in chasing the plane down a runway and is able to shoot one of the engines. Unable to take off, Castor kills the pilot and crashes the plane into a hangar. In the ensuing chaos, Pollux is captured by the FBI and Castor is knocked into a coma after bragging to Archer about a bomb that would destroy Los Angeles.
Archer believes Castor to have been bluffing, but after the FBI finds schematics for the bomb on a disk recovered from the wreckage, Archer realizes that it is real. Unable to find any information about the location of the bomb, Archer knows that the only way to obtain the location is from Castor's brother Pollux. Knowing he would never confess the bomb's location to the FBI, Archer's colleagues convince him to undergo a surgical procedure to switch faces with Castor's comatose body.
After the procedure, arrangements are made for Archer (as Castor) to be incarcerated with Pollux so that he can discover the location of the bomb which he does successfully. Castor awakens from his coma and has the procedure performed on himself, stealing Archer's face. Castor then kills everyone who knew about the operation and visits Archer in prison. Castor (as Archer) has the FBI negotiate a deal with his brother for his release in return for revealing the bomb's location. Castor disarms the bomb and revels in praise from his colleagues and the media, informing his brother that they are going straight.
Archer (as Castor) escapes from Erehwon prison, which is revealed to be inside an offshore oil platform, and swims to shore. He visits Dietrich Hassler, a known associate of Castor's and asks Dietrich for help killing Sean Archer (Castor). Archer (as Castor) meets Sasha and her son Adam, of whom Castor is the father. Pollux Troy is watching Dietrich's apartment and informs Castor of Archer's arrival. Castor (as Archer) sends an FBI team to kill Archer. Following a gunfight, Dietrich is killed by Castor (as Archer) and Archer (as Castor) manages to kill Pollux.
FBI Director Victor Lazaro berates Castor (as Archer) for the bloodbath. Castor, still angry at the loss of his brother, confesses his true identity to Lazaro and kills him, later blaming the death on a heart attack. Meanwhile, Archer (as Castor) returns to his suburban home and is able to explain the situation and convince his wife that he is really Archer.
Sasha and Archer (as Castor) track Castor to Lazaro's funeral, where Castor (as Archer) is holding Archer's wife and daughter Jamie hostage. The ensuing standoff leads to a gunfight in which Sasha and Castor's men are killed. Castor and Archer fight and Archer gains the upper hand. Jamie finds a gun and shoots at Archer, believing him to be Castor, but wounds him in the shoulder. Castor manages to escape in a boat, pursued by Archer (as Castor). After a lengthy chase both Archer and Castor's boats are destroyed and they are thrown ashore by an explosion. The two fight and Archer kills Castor with a spear gun. Archer's wife is able to convince the FBI of Sean's true identity and the face swap is reversed. The film ends with Archer adopting Sasha's son Adam.
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